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- Mud-puddling on roadsides: a potential ecological trap for butterfliesPublication . Campioni, Letizia; Marengo, Ilaria; Román, Jacinto; D'Amico, MarcelloRoad-kill represents a major threat for butterflies and more generally for pollinators. Here we report an observation of conspicuous aggregations of butterflies mud-puddling on roadsides and, for this reason, being massively road-killed by farm vehicles.